Late Ordovician cephalopods, tentaculitides, machaeridians and echinoderms from Kuh-e Faraghan, High Zagros, Iran

MOHAMMAD GHAVIDEL-SYOOKI, DAVID H. EVANS, MANSOUREH GHOBADI POUR,
LEONID E. POPOV, J. JAVIER ÁLVARO, UTKYR RAKHMONOV, INNA A. KLISHEVICH and
MOHAMMAD H. EHSANI
GHAVIDEL-SYOOKI, M., EVANS, D.H., GHOBADI POUR, M., POPOV, L.E., ÁLVARO, J.J., RAKHMONOV, U., KLISHEVICH, I.A. & EHSANI, M.D., XX.XX.2015.
Late Ordovician cephalopods, tentaculitides, machaeridians and echinoderm columnals from Kuh-e Faraghun, High Zagros, Iran. Alcheringa 39,
XXX–XXX. ISSN 0311-5518.
Late Ordovician (Katian, uppermost Acanthochitina barbata to Armoricochitina nigerica chitinozoan zones) cephalopods, tentaculitides,
machaeridians and echinoderms are documented for the first time from the southern Zagros Ranges. A low-diversity cephalopod fauna
includes Geisonocerina dargazense sp. nov., Isorthoceras sp. cf. I. bisignatum (Barrande) and other undetermined orthoceratides. The presence
of Late Ordovician tentaculitides in the high- to mid-latitude margins of Gondwana has been documented previously, but no examples have
been described in detail. Thus, Costatulites kimi sp. nov., which currently occurs associated with brachiopods characteristic of the Svobodaina
havliceki (brachiopod) Association, represents the earliest undoubted record of tentaculitides in Gondwana. Machaeridians constitute a relatively
common component of the Late Ordovician benthic faunas from the Mediterranean margin of Gondwana, but no previous records
on the Gondwanan Iranian-Arabian segment have been reported. Three echinoderm taxa based on dissociated columnals are documented from
the Armoricochitina nigerica chitinozoan Zone, including Sumsaricystis radiatus Stukalina, Ristnacrinus sp. and Rosulicrinus rosulus
Stukalina.
Mohammad Ghavidel-Syooki Institute of Petroleum Engineering, Technical Faculty of Tehran University, PO
Box 11365-4563, Tehran, Iran; David H. Evans , Natural England, Suite D, Unex House, Bourges Boulevard,
Peterborough PE1 1NG, UK; Mansoureh Ghobadi Pour* , Department of Geology, Faculty of Sciences, Golestan
University, Gorgan, Iran; Leonid E. Popov , Department of Geology, National Museum of Wales, Cathays
Park, Cardiff CF10 3NP, UK; J. Javier Álvaro , Instituto de Geociencias (CSIC-UCM), c/ José Antonio Novais 2, 28040
Madrid, Spain; Utkyr Rakhmonov, Kitab State Geological Reserve, 9 Ipak Yuli Street, Sakhrisabz, Uzbekistan; Inna A. Klishevich, Department of
Historical Geology, Geological Faculty, St Petersburg State University, Universitetskaya Naberezhnaya 7/9, 199034 St Petersburg, Russia
; Mohammad H. Ehsani , Institute of Petroleum Engineering, Technical Faculty of Tehran
University, PO Box 11365-4563, Tehran, Iran. *Also affiliated with Department of Geology, National Museum of Wales, Cathays Park, Cardiff
CF10 3NP, UK. Received 26.2.2015; revised 22.4.2015; accepted 15.5.2015.
Key words: Gondwana, Katian, Seyahou Formation, taxonomy, invertebrates, biogeography.